K. z. zeylonensis (Gmelin, 1788) – นกทึดทือพันธุ์เหนือศรีลังกา. ประเทศศรีลังกา.
Small and dark.
K. z. leschenault (Temminck, 1820) – นกทึดทือพันธุ์เหนือธรรมดาเอเชียใต้ถึงประเทศพม่า (ยกเว้นภาคตะวันออกเฉียงเหนือ) และ ประเทศไทย อาจรวมชนิด orientalis มีขนาดใหญ่กว่าและสีจางกว่า zeylonensis
K. z. semenowi (Zarudny, 1905) – นกทึดทือพันธุ์เหนือตะวันตกเลแวนต์ (อาจสูญพันธุ์แล้ว) และตอนใต้ของอานาโตเลียตลอดไปจนถึงเมโสโปเตเมีย (ข้อมูลเก่า) ถึงประเทศปากีสถาน สีซีดกว่า leschenault อาจแยกมาตั้งชนิดใหม่ได้
K. z. orientalis Delacour, 1926 – นกทึดทือพันธุ์เหนือตะวันออก ภาคตะวันออกเฉียงเหนือของประเทศพม่า, เวียดนาม และภาคตะวันออกเฉียงใต้ของจีน ยังเป็นที่สงสัยว่าต่างจาก leschenault หรือไม่ สีเข้มกว่า leschenault
Grimmett, Richard; Inskipp, Carol, Inskipp, Tim & Byers, Clive (1999) : Birds of India, Pakistan, Nepal, Bangladesh, Bhutan, Sri Lanka, and the Maldives. Princeton University Press, Princeton, N.J. ISBN 0-691-04910-6
Mlíkovský, Jiří (2002) : Cenozoic Birds of the World, Part 1: Europe. Ninox Press, Prague. แม่แบบ:Listed Invalid ISBN PDF fulltext
Mlíkovský, Jiří (2003) : Brown Fish Owl (Bubo zeylonensis) in Europe: past distribution and taxonomic status. Buteo13: 61–65. PDF fulltext
Mourer-Chauviré, Cécile (2004) : [Review of Cenozoic Birds of the World, Part 1: Europe]. Auk121 (2) : 623–627. doi:10.1642/0004-8038 (2004) 121[0623:CBOTWP]2.0.CO;2 HTML fulltext
Mourer-Chauviré, Cécile & Weesie, Peter D. M. (1986) : Bubo insularis n. sp., forme endémique insulaire de grand-duc (Aves, Strigiformes) du Pléistocène de Sardaigne et de Corse ["B. insularis, an insular endemic eagle-owl from the Pleistocene of Sardinia and Corsica"]. Revue de Paléobiologie5 (2) : 197–205 [French with English abstract]. (Mourer-Chauvire%20&%20Weesie) %201986.pdf PDF fulltext
Olsen, Jery; Wink, Michael; Sauer-Gürth, Heidi & Trost, Susan (2002) : A new Ninox owl from Sumba, Indonesia. Emu102 (3) : 223–231. doi:10.1071/MU02006 PDF fulltext
Pavia, Marco (1999) : Un cranio di Bubo insularis Mourer-Chauviré & Weesie, 1986 (Aves, Strigidae) nelle brecce ossifere del Pleistocene di Capo Figari (Sardegna, Italia) ["A cranium of B. insularis from the Pleistocene ossiferous breccia of Cape Figari (Sardinia, Italy)"]. Atti della Accademia delle Scienze di Torino, Classe di Scienze fisiche, matematiche e naturali133: 1–10 [Italian with English abstract]. PDF fulltext
Singh, A. P. (2002) : New and significant records from Dehra Dun valley, lower Garhwal Himalayas, India. Forktail18: 151–153. PDF fulltext
van den Berg, Arnoud B.; Bekir, Soner; de Knijff, Peter & The Sound Approach (2010) : Rediscovery, biology, vocalisations and taxonomy of fish owls in Turkey. Dutch Birding32: 287–298. PDF fulltext
World Owl Trust (WOT) (2005) : Brown Fish Owl. Version of February 2005. Retrieved 23 June 2009.
นกท, ดท, อพ, นธ, เหน, เส, ยงร, อง, ใช, อม, สถานะการอน, กษ, ความเส, ยงต, iucn, การจำแนกช, นทางว, ทยาศาสตร, อาณาจ, กร, animaliaไฟล, chordataช, avesอ, นด, strigiformesวงศ, strigidaeสก, ketupaสป, zeylonensisช, อทว, นามketupa, zeylonensis, gmelin, 1788, ชน, ดย, อย4. nkthudthuxphnthuehnux esiyngrxng withiich khxmul sthanakarxnurkskhwamesiyngta IUCN 3 1 1 karcaaenkchnthangwithyasastrxanackr Animaliaiflm Chordatachn Avesxndb Strigiformeswngs Strigidaeskul Ketupaspichis K zeylonensischuxthwinamKetupa zeylonensis Gmelin 1788 chnidyxy4 chnid duinbthkhwam chuxphxngStrix zeylonensis Gmelin 1788 nkthudthuxphnthuehnux xngkvs Brown fish owl chuxwithyasastr Ketupa zeylonensis hrux Bubo zeylonensis 2 epnnkekhaaemwchnidhnung phbinekhtxbxunaelaekhtrxnchuntamihlthwipexechiyaelahmuekaa 3 enuxha 1 lksna 1 1 chnidyxy 2 xangxing 3 brrnanukrm 4 aehlngkhxmulxunlksna aekikhnkthudthuxphnthuehnuxepnnkekhaaemwkhnadihyyawpraman 55 esntiemtr hnk 2 2 5 kiolkrm twphuelkkwatwemiyelknxy latwihykhlaynkekhaihy aetaekhngepluxyimmikhnpkkhlum siehluxngaekmekhiyw pakethaaekmekhiyw taehluxng hwaelakrahmxmmilaykhidsiekhm latwdanbnnatalekhmaekmethaelknxy latwdanlangnatalaekmsm milaykhidbangcakaeknkhnsidaaelamilaykhwangbangcaklaykhnsikhla khxaekmkhaw hnaphakaelaokhnpakimmikhnsikhaw esiyngrxngthumtadng hup hup hu sxngephsmilksnakhlaykn 4 chnidyxy aekikh michnidyxysamthungsichnidthiepnthiyxmrb 5 K z zeylonensis Gmelin 1788 nkthudthuxphnthuehnuxsrilngka praethssrilngka Small and dark K z leschenault 6 Temminck 1820 nkthudthuxphnthuehnuxthrrmda exechiyitthungpraethsphma ykewnphakhtawnxxkechiyngehnux aela praethsithy xacrwmchnid orientalis mikhnadihykwaaelasicangkwa zeylonensis K z semenowi Zarudny 1905 nkthudthuxphnthuehnuxtawntk elaewnt xacsuyphnthuaelw aelatxnitkhxngxanaoteliytlxdipcnthungemosopetemiy khxmuleka thungpraethspakisthan sisidkwa leschenault xacaeykmatngchnidihmid 7 K z orientalis Delacour 1926 6 nkthudthuxphnthuehnuxtawnxxk phakhtawnxxkechiyngehnuxkhxngpraethsphma ewiydnam aelaphakhtawnxxkechiyngitkhxngcin yngepnthisngsywatangcak leschenault hruxim siekhmkwa leschenaultxangxing aekikh BirdLife International 2012 Ketupa zeylonensis IUCN Red List of Threatened Species Version 2012 1 International Union for Conservation of Nature subkhnemux 16 July 2012 CS1 maint ref harv link Newton Ian Kavanagh Rodney Olsen Jerry Taylor Iain b k 2002 Ecology and Conservation of Owls Clayton VIC Australia Csiro Publishing p 346 ISBN 978 0 643 09887 9 Grimmett et al 1999 Grimmett et al 1999 WOT 2005 Mlikovsky 2003 WOT 2005 6 0 6 1 Ketupa zeylonensis rabbkhxmulkarcaaenkphnthuaebbburnakar van den Berg et al 2010 khxmmxns miphaphaelasuxekiywkb Ketupa zeylonensisbrrnanukrm aekikhGrimmett Richard Inskipp Carol Inskipp Tim amp Byers Clive 1999 Birds of India Pakistan Nepal Bangladesh Bhutan Sri Lanka and the Maldives Princeton University Press Princeton N J ISBN 0 691 04910 6 Mlikovsky Jiri 2002 Cenozoic Birds of the World Part 1 Europe Ninox Press Prague aemaebb Listed Invalid ISBN PDF fulltext Mlikovsky Jiri 2003 Brown Fish Owl Bubo zeylonensis in Europe past distribution and taxonomic status Buteo 13 61 65 PDF fulltext Mourer Chauvire Cecile 2004 Review of Cenozoic Birds of the World Part 1 Europe Auk 121 2 623 627 doi 10 1642 0004 8038 2004 121 0623 CBOTWP 2 0 CO 2 HTML fulltext Mourer Chauvire Cecile amp Weesie Peter D M 1986 Bubo insularis n sp forme endemique insulaire de grand duc Aves Strigiformes du Pleistocene de Sardaigne et de Corse B insularis an insular endemic eagle owl from the Pleistocene of Sardinia and Corsica Revue de Paleobiologie 5 2 197 205 French with English abstract Mourer Chauvire 20 amp 20Weesie 201986 pdf PDF fulltext Olsen Jery Wink Michael Sauer Gurth Heidi amp Trost Susan 2002 A new Ninox owl from Sumba Indonesia Emu 102 3 223 231 doi 10 1071 MU02006 PDF fulltext Pavia Marco 1999 Un cranio di Bubo insularis Mourer Chauvire amp Weesie 1986 Aves Strigidae nelle brecce ossifere del Pleistocene di Capo Figari Sardegna Italia A cranium of B insularis from the Pleistocene ossiferous breccia of Cape Figari Sardinia Italy Atti della Accademia delle Scienze di Torino Classe di Scienze fisiche matematiche e naturali 133 1 10 Italian with English abstract PDF fulltext Singh A P 2002 New and significant records from Dehra Dun valley lower Garhwal Himalayas India Forktail 18 151 153 PDF fulltext van den Berg Arnoud B Bekir Soner de Knijff Peter amp The Sound Approach 2010 Rediscovery biology vocalisations and taxonomy of fish owls in Turkey Dutch Birding 32 287 298 PDF fulltext World Owl Trust WOT 2005 Brown Fish Owl Version of February 2005 Retrieved 23 June 2009 khumuxdunkhmxbuysng elkhakul nkemuxngithy aelahnngsuxnkinemuxngithy elm 2 rs oxphas khxbekhttaehlngkhxmulxun aekikh khxmulekiywkhxngkb Ketupa zeylonensis cakwikispichisekhathungcak https th wikipedia org w index php title nkthudthuxphnthuehnux amp oldid 6595920, wikipedia, วิกิ หนังสือ, หนังสือ, ห้องสมุด,
